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Dr. Ziad Hafed

Organization: Werner Reichardt Centre for Integrative Neuroscience


Otfried-Müller-Straße 25
72076 Tübingen

Phone number: +49 (0)7071 29 88819

Department: CIN Physiology of Active Vision

Position: Head of Research Group

Area: CIN Members

Field of Research

Humans are tremendously reliant on the sense of vision to interact with their environment. Such interaction often involves the generation of motor outputs in response to visual stimuli, but these outputs themselves can alter the sensory inputs used by the visual system. This is particularly true of eye movements, which are necessary for high-acuity vision but at the same time cause the largest and most frequent disruptions to the retinal images projected from the eyes to the brain.
Our research aims to investigate the neural mechanisms through which visual perception interacts with motor control. We employ techniques for monitoring and focally perturbing neural activity, coupled with careful behavioral measurement and control, to understand the functional contribution of individual brain circuits in coordinating perception and action. Besides clarifying our understanding of the sense of vision, our research can also shed light on how neural activity that is distributed across multiple brain areas is organized to support behavior.


Vision, visual perception, neural mechanisms, neurophysiology, cognition, attention, microsaccades, eye movements, systems neuroscience, computational neuroscience, oculomotor system, fixational eye movements, executive function


1.            Hafed ZM, Lovejoy LP and Krauzlis RJ (2011). Modulation of microsaccades in monkey during a covert visual attention task. Journal of Neuroscience, Vol. 31, No. 43, pp. 15219-15230.
2.            Hafed ZM and Krauzlis RJ (2010). Microsaccadic suppression of visual bursts in the primate superior colliculus. Journal of Neuroscience, Vol. 30, No. 28, pp. 9542-9547.
3.            Hafed ZM, Goffart L and Krauzlis RJ (2009). A neural mechanism for microsaccade generation in the primate superior colliculus. Science, Vol. 323, No. 5916, pp. 940-943.
4.            Hafed ZM and Krauzlis RJ (2008). Goal representations dominate superior colliculus activity during extrafoveal tracking. Journal of Neuroscience, Vol. 28, No. 38, pp. 9426-9439.
5.            Hafed ZM, Goffart L and Krauzlis RJ (2008). Superior colliculus inactivation causes stable offsets in eye position during tracking. Journal of Neuroscience, Vol . 28, No. 32, pp. 8124-8137.
6.            Hafed ZM and Krauzlis RJ (2006). Ongoing eye movements constrain visual perception. Nature Neuroscience, Vol. 9, No. 11, pp. 1449-1457.